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Abstract

Results of a geomorphologic study as well as radiocarbon and pollen analyses of sediments in small basins of the Jasło-Sanok Depression (Western Carpathians) are summarised. Floors of these basins, carved in soft shale-sandstone Krosno Beds, are covered with channel fluvial deposits and oxbow-lake sediments with lake chalk and peat accumulated in the Late Vistulian and Holocene. Since the early Atlantic Phase (ca 8,400–7,900 BP) the apparent acceleration of overbank (flood) deposition intermitting the peat accumulation is observed. The plant succession includes the Late Glacial (pre-Allerød, Allerød and Younger Dryas) with coniferous park forests, through mixed deciduous forests of the Holocene with elm, hazel, oak and lime as well as spruce-elm forests with alder in wetlands, up to present-day hornbeam forests (Tilio-Carpinetum of various types) and extra-zonal Carpathian beech forests (Dentario-Glandulosae- Fagetum). Abies alba (fir) is frequent in both these association types. First evidences of synanthropic plants that prove presence of prehistoric man appeared in the Subboreal Phase. The oldest radiocarbon date 13,550±100 BP (Gd-7355) [16,710–16,085 b2k], from a bottom part of the Humniska section is probably overestimated. This is indicated by palynological data, which suggest attribution of this section to the older Allerød. Small thickness of gravel blanket from the Plenivistulian termination and the beginning of the Late Vistulian, as well as large areas devoid of weathering and solifluction covers indicate that during the Plenivistulian weathering processes and removal of silt-clay material predominated in the basins. In that time the deflation was among important processes, which is proved by deflation troughs, faceted cobbles and thick covers of the Carpathian type of loess. The Besko Basin has pre-Vistulian tectonic foundation, while landforms of its floor are of erosion-degradation origin and formed during the last Scandinavian glaciation. In the Holocene the basin floors were overbuilt with fluvial deposits up to 8 m thick.
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Abstract

The pollen morphology of many collections of taxa of the tribe Nigelleae from the family Ranunculaceae which occur worldwide is presented in this study. A total of 88 specimens from 21 taxa, some of which were recently proposed, belonging to the genera Komaroffia, Garidella, and Nigella of Nigelleae were examined using light microscopy (LM) and scanning electron microscopy (SEM). In the tribe, the pollen type is mostly trizonocolpate, but in many taxa and specimens, both trizonocolpate and non-trizonocolpate types occur together. The pollen grains are small to medium (25–53.75 μm × 20–55 μm) in size and oblate to prolate in shape. The exine pattern at the mesocolpium in all the taxa investigated is similar: micro-echinate in LM and micro-echinate-punctate in SEM. The colpus membrane in Komaroffia and Nigella is micro-echinate in both LM and SEM. In Garidella, it is micro-echinate in LM but echinate (spinulose) in SEM. In this study, multivariate analyses, principal component analysis (PCA), and unweighted pair group method with arithmetic mean (UPGMA), were used to evaluate relationships between the genera and species within the tribe with respect to pollen morphology. PCA results show three main groups in the tribe: Garidella, Komaroffia, and Nigella. Moreover, the UPGMA tree also chiefly supports generic segregation into the smaller genera. An overall synthesis of the pollen characteristics of the three genera is provided and discussed.
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